"dave:
i added them when i
"dave:
i added them when i switched from factory hx to the mercrusier versions - they were in the kit and the instructions called for their installation. As best as i can tell, you get them/omit them based on the details of the cap on top of the HX. I haven't had the stock stuff for going on 20 years so I can't tell you what they have or don't but I'm sure you can check.
On the merc caps, they are equipped like a car. they have an inner and an outer seal (or upper and lower). on the inner or center, there is a little relief valve that works (opens) when things cool off. This opening allows the fluid from the overflow connection to 'fill' the hx - it gets sucked in instead of the hoses collapsing.
When it heats up, the coolant expands and any that goes thru the relief before it closes is captured in the bottle for 'reuse'. On my big blocks, this is less than a pint. If it overheats a little, the bottle will catch it; if it is bad, the bottle usually overflows thru its vent - really no benefit in this case.
So, I'd say the tanks provide a means to ensure the HX is always full and to provide a means to inspect coolant level while keeping the system closed.
stant, the radiator cap people, used to have a great tech description on their www site but I haven't been there in a few years. I learned all this stuff by reading the manualin the radiator pressure tester kits and inspecting everything i touch."